Titelia

Fund shareholders of aTyr Pharma Inc

ISIN US0021202025 · United States · LEI 529900ENQUXMC5EBKO36

Fund shareholders
98
Of which ETFs
12 86 other funds
Value held
25.5M $
Share of capital
30.6 % of 98.1M shares on Jul 31, 2026
FundSharesValueWeight in fundShare of capital
Federated Hermes Kaufmann Fund Federated Hermes Equity Funds 5,585,0004.5M $0.10 %5.69 %as of Apr 30, 2026
Federated Hermes Kaufmann Small Cap Fund Federated Hermes Equity Funds 4,806,8773.9M $0.13 %4.90 %as of Apr 30, 2026
Fidelity Growth Company Fund Fidelity Mt. Vernon Street Trust 4,228,8324.1M $0.00 %4.31 %as of Feb 28, 2026
VANGUARD TOTAL STOCK MARKET INDEX FUND VANGUARD INDEX FUNDS 2,650,7882.1M $0.00 %2.70 %as of Mar 31, 2026
iShares Russell 2000 ETF iShares Trust 2,160,9751.7M $0.00 %2.20 %as of Mar 31, 2026
Select Biotechnology Portfolio Fidelity Select Portfolios 1,754,5941.7M $0.03 %1.79 %as of Feb 28, 2026
Fidelity Growth Company K6 Fund Fidelity Mt. Vernon Street Trust 1,358,0521.3M $0.01 %1.38 %as of Feb 28, 2026
Fidelity Series Growth Company Fund Fidelity Mt. Vernon Street Trust 1,200,3661.2M $0.01 %1.22 %as of Feb 28, 2026
VANGUARD EXTENDED MARKET INDEX FUND VANGUARD INDEX FUNDS 1,006,403785K $0.00 %1.03 %as of Mar 31, 2026
Fidelity Small Cap Index Fund Fidelity Salem Street Trust 936,184755.9K $0.00 %0.95 %as of Apr 30, 2026
iShares Russell 2000 Growth ETF iShares Trust 682,261532.2K $0.00 %0.70 %as of Mar 31, 2026
Fidelity Extended Market Index Fund Fidelity Concord Street Trust 531,354510K $0.00 %0.54 %as of Feb 28, 2026
VANGUARD RUSSELL 2000 INDEX FUND VANGUARD SCOTTSDALE FUNDS 475,772456.6K $0.00 %0.49 %as of Feb 28, 2026
iShares Micro-Cap ETF iShares Trust 237,406185.2K $0.02 %0.24 %as of Mar 31, 2026
Schwab Small-Cap Index Fund SCHWAB CAPITAL TRUST 208,534168.4K $0.00 %0.21 %as of Apr 30, 2026
Fidelity Total Market Index Fund Fidelity Concord Street Trust 168,117161.4K $0.00 %0.17 %as of Feb 28, 2026
Master Small Cap Index Series Quantitative Master Series LLC 150,707117.6K $0.00 %0.15 %as of Mar 31, 2026
iShares Core S&P Total U.S. Stock Market ETF iShares Trust 142,491111.1K $0.00 %0.15 %as of Mar 31, 2026
Fidelity Series Total Market Index Fund Fidelity Concord Street Trust 135,138129.7K $0.00 %0.14 %as of Feb 28, 2026
Federated Hermes Kaufmann Fund II Federated Hermes Insurance Series 117,80191.9K $0.10 %0.12 %as of Mar 31, 2026
Nuveen Small Cap Blend Index Fund TIAA-CREF Funds 112,97291.2K $0.00 %0.12 %as of Apr 30, 2026
EQ/2000 Managed Volatility Portfolio EQ Advisors Trust 100,39478.3K $0.00 %0.10 %as of Mar 31, 2026
VANGUARD RUSSELL 2000 GROWTH INDEX FUND VANGUARD SCOTTSDALE FUNDS 83,37780K $0.01 %0.09 %as of Feb 28, 2026
Nuveen Equity Index Fund TIAA-CREF Funds 79,39864.1K $0.00 %0.08 %as of Apr 30, 2026
Fidelity Small Cap Growth Index Fund Fidelity Salem Street Trust 67,75352.8K $0.00 %0.07 %as of Mar 31, 2026
LVIP SSGA Small-Cap Index Fund LINCOLN VARIABLE INSURANCE PRODUCTS TRUST 67,10052.3K $0.00 %0.07 %as of Mar 31, 2026
Fidelity Nasdaq Composite Index Fund Fidelity Concord Street Trust 66,66964K $0.00 %0.07 %as of Feb 28, 2026
Small Cap Index Fund Northern Funds 59,35946.3K $0.00 %0.06 %as of Mar 31, 2026
VANGUARD BALANCED INDEX FUND VANGUARD VALLEY FORGE FUNDS 56,84644.3K $0.00 %0.06 %as of Mar 31, 2026
VANGUARD INSTITUTIONAL TOTAL STOCK MARKET INDEX FUND VANGUARD INSTITUTIONAL INDEX FUNDS 45,98035.9K $0.00 %0.05 %as of Mar 31, 2026
Fidelity ZERO Total Market Index Fund Fidelity Concord Street Trust 43,13334.8K $0.00 %0.04 %as of Apr 30, 2026
Global X Russell 2000 ETF GLOBAL X FUNDS 41,53133.5K $0.00 %0.04 %as of Apr 30, 2026
Goldman Sachs Absolute Return Tracker Fund GOLDMAN SACHS TRUST 41,44932.3K $0.00 %0.04 %as of Mar 31, 2026
Small Cap Index Fund VALIC Co I 33,62032.3K $0.00 %0.03 %as of Feb 28, 2026
Fidelity ZERO Extended Market Index Fund Fidelity Concord Street Trust 31,25825.2K $0.00 %0.03 %as of Apr 30, 2026
iShares Russell 3000 ETF iShares Trust 31,00324.2K $0.00 %0.03 %as of Mar 31, 2026
ATM Small Cap Managed Volatility Portfolio EQ Advisors Trust 29,36822.9K $0.00 %0.03 %as of Mar 31, 2026
EQ/AB Small Cap Growth Portfolio EQ Advisors Trust 27,60021.5K $0.00 %0.03 %as of Mar 31, 2026
EQ/Small Company Index Portfolio EQ Advisors Trust 27,60021.5K $0.00 %0.03 %as of Mar 31, 2026
STATE STREET SMALL/MID CAP EQUITY INDEX PORTFOLIO STATE STREET INSTITUTIONAL INVESTMENT TRUST 23,70018.5K $0.00 %0.02 %as of Mar 31, 2026
Small-Cap Index Portfolio PACIFIC SELECT FUND 23,20618.1K $0.00 %0.02 %as of Mar 31, 2026
Diversified Equity Master Portfolio Master Investment Portfolio 22,62717.6K $0.00 %0.02 %as of Mar 31, 2026
1290 VT Micro Cap Portfolio EQ Advisors Trust 17,19913.4K $0.01 %0.02 %as of Mar 31, 2026
NVIT Small Cap Index Fund Nationwide Variable Insurance Trust 16,48312.9K $0.00 %0.02 %as of Mar 31, 2026
Bridge Builder Small/Mid Cap Growth Fund Bridge Builder Trust 15,89712.4K $0.00 %0.02 %as of Mar 31, 2026
Fidelity Flex Small Cap Index Fund Fidelity Salem Street Trust 15,76512.7K $0.00 %0.02 %as of Apr 30, 2026
iShares Russell Small/Mid-Cap Index Fund BlackRock Funds 13,66311K $0.00 %0.01 %as of Apr 30, 2026
DWS Small Cap Index VIP DEUTSCHE DWS INVESTMENTS VIT FUNDS 13,31110.4K $0.00 %0.01 %as of Mar 31, 2026
EQ/Morgan Stanley Small Cap Growth Portfolio EQ Advisors Trust 13,15510.3K $0.00 %0.01 %as of Mar 31, 2026
SIIT EXTENDED MARKET INDEX FUND SEI INSTITUTIONAL INVESTMENTS TRUST 13,00012.5K $0.00 %0.01 %as of Feb 28, 2026
BlackRock Small Cap Index V.I. Fund BlackRock Variable Series Funds, Inc. 11,0248.6K $0.00 %0.01 %as of Mar 31, 2026
Blackstone Alternative Multi-Strategy Fund Blackstone Alternative Investment Funds 10,8008.4K $0.00 %0.01 %as of Mar 31, 2026
Voya RussellTM Small Cap Index Portfolio Voya VARIABLE PORTFOLIOS INC 10,3388.1K $0.00 %0.01 %as of Mar 31, 2026
SA Small Cap Index Portfolio SUNAMERICA SERIES TRUST 10,1508.2K $0.00 %0.01 %as of Apr 30, 2026
CVT Russell 2000 Small Cap Index Portfolio Calvert Variable Trust, Inc. 8,8816.9K $0.00 %0.01 %as of Mar 31, 2026
PD Small-Cap Growth Index Portfolio PACIFIC SELECT FUND 8,7166.8K $0.01 %0.01 %as of Mar 31, 2026
T. Rowe Price Small-Cap Index Fund T. ROWE PRICE INDEX TRUST, INC. 8,5186.6K $0.00 %0.01 %as of Mar 31, 2026
Strategic Equity Allocation Trust John Hancock Variable Insurance Trust 8,2496.4K $0.00 %0.01 %as of Mar 31, 2026
Nationwide Small Cap Index Fund Nationwide Mutual Funds 7,6426.2K $0.00 %0.01 %as of Apr 30, 2026
iShares Total U.S. Stock Market Index Fund BlackRock Funds 7,4876K $0.00 %0.01 %as of Apr 30, 2026
VANGUARD RUSSELL 3000 INDEX FUND VANGUARD SCOTTSDALE FUNDS 7,3667.1K $0.00 %0.01 %as of Feb 28, 2026
Venerable Small Cap Index Fund Venerable Variable Insurance Trust 7,2055.6K $0.00 %0.01 %as of Mar 31, 2026
SmallCap Growth Fund I Principal Funds, Inc 6,7095.4K $0.00 %0.01 %as of Apr 30, 2026
Fidelity Small Cap Value Index Fund Fidelity Salem Street Trust 6,3174.9K $0.00 %0.01 %as of Mar 31, 2026
VIP Extended Market Index Portfolio Variable Insurance Products Fund II 6,1124.8K $0.00 %0.01 %as of Mar 31, 2026
ProShares Ultra Russell2000 ProShares Trust 5,9365.7K $0.00 %0.01 %as of Feb 28, 2026
ProShares UltraPro Russell2000 ProShares Trust 5,9155.7K $0.00 %0.01 %as of Feb 28, 2026
Morgan Stanley Pathway Small-Mid Cap Equity ETF MORGAN STANLEY PATHWAY FUNDS 4,7884.6K $0.00 %0.00 %as of Feb 28, 2026
AST Small-Cap Equity Portfolio Advanced Series Trust 4,5403.5K $0.00 %0.00 %as of Mar 31, 2026
VANGUARD RUSSELL 2000 VALUE INDEX FUND VANGUARD SCOTTSDALE FUNDS 4,3414.2K $0.00 %0.00 %as of Feb 28, 2026
ProShares Russell 2000 High Income ETF ProShares Trust 4,1544K $0.00 %0.00 %as of Feb 28, 2026
JNL/DFA U.S. Small Cap Fund JNL Series Trust 4,1453.2K $0.00 %0.00 %as of Mar 31, 2026
State Street Balanced Index Fund STATE STREET INSTITUTIONAL INVESTMENT TRUST 3,3932.6K $0.00 %0.00 %as of Mar 31, 2026
State Street Total Return V.I.S. Fund STATE STREET VARIABLE INSURANCE SERIES FUNDS INC 3,3322.6K $0.00 %0.00 %as of Mar 31, 2026
Bridge Builder Small/Mid Cap Value Fund Bridge Builder Trust 3,3072.6K $0.00 %as of Mar 31, 2026
EQ/Franklin Small Cap Value Managed Volatility Portfolio EQ Advisors Trust 3,1402.4K $0.00 %0.00 %as of Mar 31, 2026
Dimensional U.S. Core Equity 2 ETF Dimensional ETF Trust 3,1262.5K $0.00 %as of Apr 30, 2026
Penn Series Small Cap Index Fund Penn Series Funds Inc 2,8002.2K $0.00 %0.00 %as of Mar 31, 2026
Destinations Small-Mid Cap Equity Fund Brinker Capital Destinations Trust 2,6102.5K $0.00 %0.00 %as of Feb 28, 2026
Nationwide Fundamental All Cap Equity Portfolio Nationwide Mutual Funds 2,5962.1K $0.00 %0.00 %as of Apr 30, 2026
Total Stock Market Index Trust John Hancock Variable Insurance Trust 2,1831.7K $0.00 %0.00 %as of Mar 31, 2026
VIP Total Market Index Portfolio Variable Insurance Products Fund II 2,0511.6K $0.00 %0.00 %as of Mar 31, 2026
TIAA Separate Account VA-1 - Stock Index Account TIAA Separate Account VA-1 1,9101.5K $0.00 %0.00 %as of Mar 31, 2026
Nuveen Life Stock Index Fund TIAA-CREF Life Funds 1,175916.5 $0.00 %0.00 %as of Mar 31, 2026
1290 VT Small Cap Value Portfolio EQ Advisors Trust 1,109865 $0.00 %0.00 %as of Mar 31, 2026
ULTRA SMALL CAP PROFUND ProFunds 1,084875.2 $0.00 %0.00 %as of Apr 30, 2026
SmallCap Value Fund II Principal Funds, Inc 620500.6 $0.00 %as of Apr 30, 2026
Knights of Columbus U.S. ALL CAP INDEX FUND ADVISORS' INNER CIRCLE III 507409.4 $0.00 %0.00 %as of Apr 30, 2026
Russell 2000 2x Strategy Fund Rydex Dynamic Funds 444346.3 $0.00 %0.00 %as of Mar 31, 2026
PROFUND VP ULTRA SMALL CAP ProFunds 321250.4 $0.00 %0.00 %as of Mar 31, 2026
PROFUND VP SMALL CAP ProFunds 242188.8 $0.00 %0.00 %as of Mar 31, 2026
Russell 2000 Fund Rydex Series Funds 203158.3 $0.00 %0.00 %as of Mar 31, 2026
SMALL-CAP PROFUND ProFunds 175141.3 $0.00 %0.00 %as of Apr 30, 2026
Russell 2000 1.5x Strategy Fund Rydex Series Funds 12194.4 $0.00 %0.00 %as of Mar 31, 2026
ProShares Hedge Replication ETF ProShares Trust 9894.1 $0.00 %0.00 %as of Feb 28, 2026
Columbia Research Enhanced Small Cap ETF Columbia ETF Trust I 9574.1 $0.00 %0.00 %as of Mar 31, 2026
ProShares Russell 2000 Dynamic Buffer ETF ProShares Trust 2927.8 $0.00 %0.00 %as of Feb 28, 2026
Russell 2000(R) 1.5x Strategy Fund Rydex Variable Trust 2317.9 $0.00 %0.00 %as of Mar 31, 2026

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The funds most exposed to it

The weight is the share of a fund's net assets. The holders table counts shares; this one measures what the company weighs inside each fund.

FundWeight in fund
Federated Hermes Kaufmann Small Cap Fund Federated Hermes Equity Funds 0.13 %as of Apr 30, 2026
Federated Hermes Kaufmann Fund II Federated Hermes Insurance Series 0.10 %as of Mar 31, 2026
Federated Hermes Kaufmann Fund Federated Hermes Equity Funds 0.10 %as of Apr 30, 2026
Select Biotechnology Portfolio Fidelity Select Portfolios 0.03 %as of Feb 28, 2026
iShares Micro-Cap ETF iShares Trust 0.02 %as of Mar 31, 2026
1290 VT Micro Cap Portfolio EQ Advisors Trust 0.01 %as of Mar 31, 2026
PD Small-Cap Growth Index Portfolio PACIFIC SELECT FUND 0.01 %as of Mar 31, 2026
Fidelity Growth Company K6 Fund Fidelity Mt. Vernon Street Trust 0.01 %as of Feb 28, 2026
VANGUARD RUSSELL 2000 GROWTH INDEX FUND VANGUARD SCOTTSDALE FUNDS 0.01 %as of Feb 28, 2026
Fidelity Series Growth Company Fund Fidelity Mt. Vernon Street Trust 0.01 %as of Feb 28, 2026

Quarter after quarter

Each SEC quarterly data set, newest first. The change is read against the quarter before it.

QuarterShareholdersChangeSharesChange
2026Q298-229,968,095-0.4 %
2026Q1100-1330,097,690-9.7 %
2025Q4113+2233,329,719+0.7 %
2025Q391+5733,088,406+19.3 %
2025Q234027,731,159+4.1 %
2025Q134+726,641,657+2.9 %
2024Q427025,891,663+4.4 %
2024Q327-124,792,221+27.6 %
2024Q228+519,432,183+1.6 %
2024Q123-319,132,838+0.6 %
2023Q42619,014,905

Institutional managers

107 managers declare this company to the SEC (Form 13F); the 25 largest follow.

ManagerSharesValue
FMR LLC14,706,84311.5M $as of Mar 31, 2026
FEDERATED HERMES, INC.10,509,6788.2M $as of Mar 31, 2026
BlackRock, Inc.6,443,9045M $as of Mar 31, 2026
UBS Group AG3,867,2133M $as of Mar 31, 2026
GEODE CAPITAL MANAGEMENT, LLC2,385,4851.9M $as of Mar 31, 2026
STATE STREET CORP1,754,3991.4M $as of Mar 31, 2026
GSA CAPITAL PARTNERS LLP1,684,5861.3K $as of Mar 31, 2026
MILLENNIUM MANAGEMENT LLC1,641,2461.3M $as of Mar 31, 2026
MORGAN STANLEY1,225,467955.9K $as of Mar 31, 2026
TWO SIGMA INVESTMENTS, LP1,144,565892.8K $as of Mar 31, 2026
SUSQUEHANNA INTERNATIONAL GROUP, LLP866,744676.1K $as of Mar 31, 2026
NORTHERN TRUST CORP711,105554.7K $as of Mar 31, 2026
AQR CAPITAL MANAGEMENT LLC496,018386.9K $as of Mar 31, 2026
BANK OF AMERICA CORP /DE/303,559236.8K $as of Mar 31, 2026
GOLDMAN SACHS GROUP INC283,236220.9K $as of Mar 31, 2026
Nuveen, LLC229,284178.8K $as of Mar 31, 2026
CHARLES SCHWAB INVESTMENT MANAGEMENT INC210,737164.4K $as of Mar 31, 2026
Bank of New York Mellon Corp193,531151K $as of Mar 31, 2026
GROUP ONE TRADING LLC184,014143.5K $as of Mar 31, 2026
MARSHALL WACE, LLP138,118107.7K $as of Mar 31, 2026
XTX Topco Ltd111,42386.9K $as of Mar 31, 2026
DEUTSCHE BANK AG\103,32680.6K $as of Mar 31, 2026
RHUMBLINE ADVISERS103,14680.4K $as of Mar 31, 2026
HOHIMER WEALTH MANAGEMENT, LLC100,00078K $as of Mar 31, 2026
DIMENSIONAL FUND ADVISORS LP91,87171.5K $as of Mar 31, 2026

Declared shareholders of aTyr Pharma Inc

1 beneficial ownership filing. These stakes do not add up: an asset manager filing aggregates its own funds, already counted above.

ShareholderDeclared stakeShares
FMR LLC and 1 other6.10 %5,935,900as of May 29, 2026 · SCHEDULE 13G
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Titelia. "Fund shareholders of aTyr Pharma Inc". https://titelia.com/en/stocks/atyr-pharma-inc-US0021202025